I have a sub assembly that is fully constrained.  It is up to date, with no warnings or errors.  There is one configuration and one display state.  

When I open this subassembly in the parent assembly, the parts are in the wrong position.  (The "lock rotation" constraint is applied 180 degrees off.)

I've had this problem off and on since 2018, although the symptoms are sometimes different. 

Any ideas on what I'm doing wrong, or how to fix it?  

Sub assembly:

Parent assembly:

The parts in question are located with a concentric mate, with lock rotation turned on.